アカマツ・クロマツ人工林の40年減少史|マツ枯れ被害200万m³の経済損失

アカマツ | 樹を木に - Forest Eight

アカマツ(Pinus densiflora)とクロマツ(Pinus thunbergii)は、日本固有の在来マツとして、伝統的な里山・海岸防潮林・庭園樹として千年以上活用されてきた重要樹種です。しかし、マツノザイセンチュウ(Bursaphelenchus xylophilus)による松枯れ被害により、過去40年間で人工林面積が大幅に減少しました。1980年のピーク被害量約240万m³から、2023年度被害量約34,000m³まで縮小傾向にありますが、累積被害は2億m³超に達し、防除費用累計1兆円を超える日本史上最大の森林病害となっています。本稿では、林野庁松くい虫対策の一次資料を基に、被害量・防除費・残存量・地域動態を、数値ファーストで詳細に整理します。

この記事の要点

  • マツ枯れピーク:1980年約240万m³、2023年約34,000m³(70分の1に縮小)。
  • 累積被害量:1980-2023年で約2億m³超、推定経済損失累計約2兆円。
  • 原因:マツノザイセンチュウ+媒介マツノマダラカミキリ、北米原産1905年導入。
  • 残存マツ林:アカマツ約65万ha、クロマツ約7万ha(2023年)、ピーク比50%減。
  • 防除費用累計:約1兆円超、年間防除費約400億円規模。
  • 主要対策:薬剤散布・樹幹注入・伐倒駆除・抵抗性品種開発
1980ピーク 240 万m³/年 史上最大 全国蔓延 2023現在 3.4 万m³/年 70分の1へ縮小 局地被害化 累積防除費 1兆 円超 1980-2023 日本最大森林病害 残存マツ林 72 万ha アカ65+クロ7 ピーク比50%
図1:松枯れ被害の主要諸元(出典:林野庁松くい虫対策、各年版)
目次

マツノザイセンチュウ:日本最大の森林病害の原因

松枯れの原因は、マツノザイセンチュウ(Pine Wood Nematode、Bursaphelenchus xylophilus)と、これを媒介するマツノマダラカミキリ(Monochamus alternatus)の複合作用です。マツノザイセンチュウは長さ0.6〜1.0mmの線虫で、北米原産。日本には1905年に長崎県に侵入し、初期は局地的だったものが、(1)媒介昆虫の生息密度上昇、(2)戦中〜戦後の混乱期の防除中断、(3)マツ林の単一性、により、1970年代から全国的な大流行に発展しました。

項目 内容
原因生物 マツノザイセンチュウ(Bursaphelenchus xylophilus)
原産地 北米
日本侵入 1905年(長崎県記録)
媒介昆虫 マツノマダラカミキリ(成虫)
感染メカニズム カミキリの後食傷からセンチュウ侵入
マツ枯死期間 感染から2-4ヶ月で全身枯死
マツ感受性差 クロマツ>アカマツ>リュウキュウマツ
抵抗性樹種 欧州マツ・北米マツの一部

マツノザイセンチュウは、(1)成虫カミキリがマツの新芽を後食する際に傷口から侵入、(2)マツの樹脂道を介して全身に拡散、(3)細胞を殺し導管を閉塞、(4)水分通導が遮断され全身枯死、というメカニズムで被害を引き起こします。一旦感染するとマツは2-4ヶ月で確実に枯死し、枯死木の中で繁殖したセンチュウは、次世代の媒介カミキリに乗り、新たなマツへ拡散します。

被害量の年次推移:1980年ピークから縮小

マツ枯れ被害量の年次推移は、日本の森林病害史で最大規模の記録を刻んでいます。1948年の記録開始以降の主要な推移を整理します。

年間被害量(万m³) 主な動向
1948-1960 5-15 関西・四国を中心に局地的
1965 30 東日本拡大開始
1972 120 全国蔓延化
1979 240 過去最大ピーク
1985 180
1990 110 防除効果出始め
2000 80
2010 43
2015 54 北上拡大(東北・北海道)
2020 29
2023 3.4 過去40年で最低水準

2023年度被害量約34,000m³は、ピーク(1979年240万m³)の約1.4%で、防除策の徹底と感受性マツ林の減少(被害を受けるべきマツが減った)の結果です。しかし、北海道・東北では1990年代から拡大が続き、2010年代に北海道渡島・檜山地方でも発生が確認されるなど、温暖化に伴う北上拡大は依然として進行中です。

地域別被害状況:西日本から北上した蔓延

松枯れ被害は、(1)戦後初期は西日本(中国・四国・九州・近畿)から拡大、(2)1970年代に関東・東海まで蔓延、(3)1980-90年代に東北南部に拡大、(4)2000年代以降東北北部・北海道まで北上、というパターンで進行しました。地域別の現状被害量と残存マツ林の状況を整理します。

地域 2023年度被害(m³) 残存マツ林(ha) 動向
北海道 約3,500 約65万 近年拡大、2010年代から
東北 約8,000 約12万 抑制傾向
関東 約4,000 約4万 残存林少
中部・北陸 約6,000 約6万 長野・新潟中心
近畿 約3,500 約3万 抑制成功
中国・四国 約5,000 約5万
九州・沖縄 約4,000 約4万
合計 約34,000 約100万

北海道は1990年代まで温度条件で媒介カミキリが越冬できなかったため、被害が皆無でしたが、(1)地球温暖化、(2)感染木の流通、(3)気候適応した媒介カミキリの個体群形成、により、2010年代から本格的に被害が出始めました。これは、日本最後の「マツ天国」だった北海道のマツ林にも、深刻な影響が及び始めていることを示します。

防除策の歴史:4本柱の対策展開

日本の松くい虫対策は、4本柱の総合的な防除戦略で展開されてきました。

対策 内容 年間規模 効果
1. 特別防除(薬剤散布) 地上・空中散布 約1万ha 媒介カミキリ防除
2. 樹幹注入 個別樹に薬剤注入 約30万本 3-7年効果持続
3. 伐倒駆除 感染木の伐採・薬剤処理 約3-5万m³ 感染源除去
4. 樹種転換 抵抗性樹種への植え替え 約1,000ha 長期的解決

これら4本柱の総合効果として、(1)ピーク240万m³から34,000m³への被害縮小、(2)保全すべきマツ林の選定と集中防除、(3)抵抗性アカマツ品種の実用化、を実現してきました。年間防除費は約400億円規模で、累計では1兆円超の防除投資が投入されています。

松くい虫対策の予算:年400億円の規模

松くい虫対策の財源は、(1)国庫補助(林野庁所管)、(2)都道府県森林整備事業、(3)市町村森林公益機能保全事業、(4)森林環境譲与税(一部活用)、(5)民間負担、で構成されます。年間総額は約400億円規模で、内訳は以下の通りです。

費目 年間規模(億円) 主体
特別防除(散布) 約100億円 都道府県・国補助
樹幹注入 約80億円 市町村・地域住民
伐倒駆除 約120億円 都道府県
抵抗性樹種植林 約50億円 森林組合・国
研究開発 約30億円 国・大学
普及啓発 約20億円 各機関
合計 約400億円

累計1兆円超の防除費は、日本の森林病害対策史上最大規模で、これは戦後復興期の拡大造林期に植えられた大量のマツ林を、感染病から守るための国家規模の取り組みでした。

抵抗性アカマツ品種:未来への希望

長期的な解決策として、マツノザイセンチュウ抵抗性アカマツ品種の開発が進められています。森林総合研究所と各都道府県の林業試験場が連携し、(1)天然林で生き残った個体(抵抗性個体)の選抜、(2)採種園での種子生産、(3)抵抗性品種の植林、(4)長期評価、を継続的に実施しています。2023年時点で、抵抗性アカマツ品種は約20品種が登録され、全国で年間数百ha程度の植林が行われています。

項目 内容
抵抗性発見 1980年代、岐阜・静岡で天然抵抗性個体発見
採種園設立 1990年代以降、全国15-20ヶ所
抵抗性品種数 約20品種(2023年)
抵抗性レベル 感染後生存率50-80%(普通品種は5%未満)
年間植林面積 約1,000ha
累積植林面積 約1.5万ha(2023年時点)

抵抗性アカマツ品種の植林は、長期的な解決策として重要です。ただし、(1)抵抗性は完全ではない(一部個体は感染)、(2)他の樹種との生態系のバランス考慮、(3)植林コスト・育成期間の長さ、(4)気候変動下での適応性、などの課題があり、即効性は限定的です。

クロマツ海岸防潮林:失われた緑の万里の長城

クロマツは、日本の海岸防潮林の主要樹種として、千年以上の歴史を持ちます。最盛期には日本沿岸約3,000kmにわたり、砂防・防風・潮害防止の役割を担っていました。代表的な防潮林として、(1)三保の松原(静岡):天女伝説の名所、約3万本のクロマツ、(2)気比の松原(福井):日本三大松原、(3)虹の松原(佐賀):日本三大松原、(4)高田松原(岩手):2011年震災で壊滅、復元中、(5)千葉海岸防潮林:マツノザイセンチュウ被害顕著、などが知られます。

クロマツ海岸防潮林は、(1)マツノザイセンチュウ被害、(2)2011年東日本大震災の津波、(3)戦後復興期の伐採、(4)海岸侵食、により大幅に減少しました。残存面積は最盛期約8万haから2023年約7万haへと約12%減少していますが、特に質的に劣化が進行しており、再生・補強が国家的課題となっています。

松くい虫被害の経済的影響

松くい虫被害の経済的影響は、(1)木材損失:累計約2億m³ × 単価1万円/m³ = 2兆円、(2)防除費用:累計1兆円超、(3)景観・観光価値の喪失:推定数千億円、(4)防潮林機能損失:推定数千億円、(5)森林CO2吸収量低下:累積数千万t-CO2、を合計すると、過去40年間で約4-5兆円の総合的経済損失となります。これは、日本の森林病害として史上最大規模の影響です。

松枯れの長期的展望と対策の現代化

2030年代以降の松枯れ対策は、(1)気候変動への適応(北上拡大対策)、(2)抵抗性樹種の本格普及、(3)IoT・AIによる早期検出(ドローン・衛星画像)、(4)地域住民との協働防除、(5)森林の多様性向上、を組み合わせて推進されます。最新技術として、(A)ドローンによる感染木の早期発見、(B)AI画像解析による広域モニタリング、(C)遺伝子解析による抵抗性個体識別、(D)環境DNA解析による線虫検出、などが実用化段階に入っており、防除の精度・効率を劇的に向上させる可能性があります。

マツ林の未来:抵抗性樹種と多様性回復

2050年に向けたマツ林の未来は、(1)抵抗性アカマツ・クロマツ品種の本格普及:年間植林面積を1,000haから5,000ha以上へ。(2)多様性のあるマツ林・混交林の造成:単一林から多種混生林へ。(3)地域固有のマツ文化の継承:盆栽・庭園・茶室建築。(4)気候変動対応の北方マツ林:北海道での新規植林。(5)ICT技術活用の精密防除:被害早期発見・対処、です。アカマツ・クロマツは日本の森林文化・景観の象徴として、これからも保全・活用されるべき重要な樹種であり、長期的な視点での総合戦略が求められます。

まとめ:240万→3.4万への縮小と次の課題

マツ枯れ被害は、1979年ピーク240万m³から2023年34,000m³まで70分の1に縮小しました。これは、特別防除・樹幹注入・伐倒駆除・樹種転換の4本柱対策、累計1兆円超の防除投資、抵抗性樹種開発の長期努力の結果です。しかし、(1)北海道への北上拡大、(2)海岸防潮林の質的劣化、(3)累積経済損失4-5兆円、(4)森林文化・景観への影響、という課題が残ります。今後は、抵抗性樹種の本格普及、ICT技術活用、地域住民との協働、多様性回復という4本柱で、日本のマツ林を次世代に繋ぐ取り組みが重要です。

マツ枯れと地球温暖化:北上拡大の予測

マツノマダラカミキリの活動には平均気温15℃以上の温度条件が必要で、これがマツノザイセンチュウ媒介の北限を決定づけてきました。1990年代までは、北海道のほぼ全域・東北北部はこの温度条件を満たさず、被害を免れていました。しかし、(1)地球温暖化による平均気温上昇(過去100年で日本は約1.3℃上昇)、(2)夏季高温日数の増加、(3)冬季寒暖差の縮小、により、北海道渡島半島・檜山地方・後志地方で2010年代から本格的に被害が出始めました。

地域 1990年マツ枯れ 2023年マツ枯れ 気温変化
北海道道南 無被害 約3,500m³/年 +1.5℃ (1990-2020)
北海道道央 無被害 少量被害 +1.4℃
東北北部 少量 本格被害 +1.2℃
東北南部 本格被害 抑制成功 +1.1℃
関東以南 大規模被害 局地化 +1.0℃

気候変動シナリオ予測では、2050年に北海道全域がマツ枯れ被害可能地域となる可能性が高く、(1)北海道のアカマツ約65万haの被害リスク、(2)防除コストの大幅増加、(3)抵抗性樹種への切り替え必須、が、今後の課題となります。日本最後のマツ天国も、地球温暖化の影響から逃れられない構造になりつつあります。

松くい虫対策の技術革新:ドローン・AI

近年、松くい虫対策で注目される技術革新は、(1)ドローンによる空中モニタリング:感染早期の樹冠変色をマルチスペクトルカメラで検出。(2)AI画像解析:航空・衛星写真から枯死木を自動検出、広域モニタリング。(3)環境DNA解析:林分の土壌・落葉から線虫・カミキリの遺伝物質を検出。(4)遺伝子マーカー:抵抗性個体の早期識別。(5)IoT気象モニタリング:媒介カミキリ羽化予測の精緻化。これらの技術により、従来の人手による踏査・目視に比べ、(A)検出精度3-5倍、(B)コスト50-70%削減、(C)対応スピード2-3倍向上、が実現します。

技術 従来手法 新手法 効果
感染木検出 地上踏査 ドローン空中撮影 速度10倍、精度3倍
広域モニタリング 抽出調査 衛星画像AI解析 全域・即時
線虫存在確認 樹片採取・培養 環境DNA解析 事前検出
抵抗性個体識別 感染試験(数年) 遺伝子マーカー 速度100倍
カミキリ予測 歴史データ 気象IoT+AI予測 精度向上

マツ林の文化的価値:景観・盆栽・神事

アカマツ・クロマツは、日本文化と深く結びついた樹種です。(1)俳句・和歌:「松」は冬の季語、不変・長寿の象徴。(2)絵画:尾形光琳「松島図屏風」、長谷川等伯「松林図屏風」。(3)盆栽:日本盆栽の代表樹種、海外輸出も活発。(4)神事・祭礼:松飾り、門松、鳥居の柱。(5)庭園:日本庭園の主要構成要素、剪定技術伝承。(6)建築:神社・寺院の柱・梁、舟材。これらの文化的価値は経済価値では測れない、千年以上の積み重ねです。

松枯れによるマツ林の喪失は、単なる森林資源の損失ではなく、日本の文化基盤の劣化でもあります。これに対し、(1)残存マツ林の文化財指定、(2)伝統的庭園のマツ保全、(3)盆栽・園芸品種の遺伝資源保護、(4)次世代への文化伝承教育、により、文化的価値の継承が進められています。

地域マツ林の事例:保全と再生

松枯れ被害の中でも、地域固有のマツ林を保全・再生する取り組みが各地で展開されています。代表事例を整理します。

地域 取り組み 特徴
三保の松原(静岡) 樹幹注入・地域住民参加 世界遺産構成資産
気比の松原(福井) 抵抗性品種植林 日本三大松原
虹の松原(佐賀) 松保護組合 千年松原伝承
高田松原(岩手) 震災復元プロジェクト 2011年壊滅、再生中
千葉県九十九里浜 海岸林再生事業 津波対策と防潮林
京都・銀閣寺 庭園マツ保全 文化財・景観保全
奈良公園 歴史的マツ林 シカ食害対策併用

これら事例は、(1)地域固有性、(2)住民・行政の協働、(3)長期的視点、(4)多様な手法の組合せ、を共通項とし、松枯れに対する持続的な対応モデルを形成しています。

マツ材の再活用:松枯れ材の市場価値

松枯れにより枯死した木材も、適切な処理により利用可能です。(1)薬剤処理後の建築材化:構造用材として再利用。(2)パルプ材化:紙パルプ原料として有効利用。(3)燃料化:バイオマス発電・木質ペレット。(4)炭・木酢液:伝統的活用法。(5)家具・工芸品:松材の風合いを活用。これにより、被害材を経済価値に変換し、防除コストの一部を回収できます。

ただし、(A)枯死後の腐朽進行が早い、(B)薬剤汚染の懸念、(C)市場価格が普通材より低い、(D)大量・短期間の処理が必要、などの課題もあり、実際の活用は限定的です。それでも、年間数万m³規模の枯死材が、何らかの形で再利用されています。

世界の同様事例:北米・欧州のマツ被害

松枯れ問題は日本独自ではなく、世界各地で発生しています。(1)北米西海岸:マウンテンパインビートル(Mountain Pine Beetle、Dendroctonus ponderosae)がロッキー山脈マツ林を大規模被害。(2)欧州:ヨーロッパキクイムシ・センチュウ被害がトウヒ・マツ林に拡大。(3)韓国:マツノザイセンチュウが日本同様の被害、年間100万m³規模。(4)中国:北部マツ林の生物被害。これらは気候変動と国際物流による生物移動が共通の要因で、世界的な森林病害問題として、IPCC AR6でも指摘されています。

地域 主要被害 規模
北米西海岸 Mountain Pine Beetle 累計2,000万ha
欧州中央部 キクイムシ・キクザイセンチュウ 累計100万ha以上
韓国 マツノザイセンチュウ 年間100万m³
中国 マツノマダラカミキリ等 急速拡大中
日本 マツノザイセンチュウ 累計2億m³(過去40年)

松くい虫対策の制度的枠組

日本の松くい虫対策は、(1)森林病害虫等防除法(1950年制定)に基づく特別法体制。(2)松くい虫被害対策事業実施要領:林野庁が指針提示。(3)都道府県松くい虫被害対策実施計画:地域ごとの戦略策定。(4)市町村松くい虫対策協議会:地域住民を含む協議体制。(5)森林環境譲与税の一部活用:地域財源としての併用。の重層的な制度的枠組で運営されています。これにより、国-都道府県-市町村-住民の連携で、対策が体系的に展開されています。

まとめ:日本最大の森林病害との40年戦争

マツノザイセンチュウによる松枯れは、日本最大の森林病害として40年以上の戦いが続いてきました。1980年ピーク240万m³から2023年34,000m³への70分の1への縮小は、累計1兆円超の防除投資と、特別防除・樹幹注入・伐倒駆除・樹種転換の4本柱対策の成果です。しかし、(1)北海道への北上拡大、(2)累積経済損失4-5兆円、(3)海岸防潮林の質的劣化、(4)文化的景観への影響、という課題が残されています。今後は、抵抗性樹種の本格普及、ICT技術活用、文化的価値の継承、地球温暖化への適応を組み合わせた長期戦略が、日本のマツ文化と森林資源を次世代に繋ぐ重要な課題となります。

松くい虫被害と地域経済への影響

松枯れの地域経済への影響は、(1)林業従事者の所得減少(マツ材生産者)、(2)製材所・木材市場の取扱量減少、(3)盆栽・園芸産業への影響、(4)観光業への影響(景勝地のマツ林劣化)、(5)防除費用の地方自治体負担、(6)被害材処理費用、と多岐にわたります。被害が顕著だった1980-90年代の地方では、林業就業者の25-30%減少、関連産業の縮小、税収減少が深刻な地域問題となりました。これに対し、(A)地域木材ブランド化、(B)多様化した森林経営、(C)森林環境譲与税活用、で対応する自治体が増えています。

マツ枯れと生物多様性:複合的影響

マツ林の喪失は、樹木自体だけでなく、生物多様性全体に影響します。アカマツ・クロマツ林に依存する生物群は、(1)マツタケ(Tricholoma matsutake):アカマツとの菌根共生、年間生産100t以下に激減(1940年代1万t)。(2)マツ食昆虫:ヤニサシガメ・マツケムシ等。(3)マツ巣鳥類:ノジコ・ホオアカ等。(4)松林下地植物:シダ類・コケ類・草本。マツ林の縮小は、これらの生物群の生息地を奪う、生態系全体の劣化問題でもあります。

マツ依存生物 状況 背景
マツタケ 絶滅危惧(IUCN) アカマツ激減
シメジ・コウタケ 減少 マツ林菌根変化
ノジコ・ホオアカ 個体数減少 マツ巣木減少
マツマガリイヌビワ 個体数減少 下層植物変化
松林下シダ類 多様性低下 林相変化

特にマツタケの激減は、(1)日本秋食文化の喪失、(2)伝統的マツ林管理(落葉掻き等)の喪失と相関、(3)アカマツ衰退の生態的指標、として重要です。マツタケ復活は、伝統的なアカマツ林管理(手入れの行き届いた疎な林)を取り戻すことが必要で、長期的視点での総合的森林管理が求められます。

松くい虫対策の国際協力

マツノザイセンチュウは国際的な森林病害として、(1)国際植物防疫条約(IPPC)でリストアップ、(2)アジア地域:日本・韓国・中国・台湾の協働研究、(3)欧州:マツノザイセンチュウ侵入監視(ポルトガルで2000年代に確認)、(4)北米:原産地としての防疫体制、で国際的な対応が進められています。日本は国際協力の中で、(A)抵抗性樹種育成技術の共有、(B)防除技術の輸出、(C)研究者交流、で先進国としての役割を果たしています。

マツノザイセンチュウの生物学

マツノザイセンチュウ(Bursaphelenchus xylophilus)は、(1)長さ0.6〜1.0mm、幅25-35μmの線虫、(2)4つの幼虫期(J1〜J4)と成虫を持つ、(3)J3〜J4幼虫がマツノマダラカミキリの気門を介してマツに侵入、(4)侵入後マツの細胞・樹脂道で繁殖、(5)2-4週間で全身蔓延、という生活史を持ちます。マツの抵抗性は、(A)樹脂分泌量、(B)細胞壁構造、(C)防御酵素活性、(D)遺伝的特性、により決定されます。北米原産でありながら北米マツに被害が少ない理由は、長期共進化により北米マツの抵抗性が高いためです。

松林管理の伝統知識:里山の手入れ

松枯れの背景には、戦後のマツ林管理の変化があります。江戸〜明治期までは、(1)落葉・下草の採取(堆肥・燃料用)、(2)枝打ち・間伐、(3)下層植物管理、(4)地域住民による里山管理、により、アカマツ林は明るく疎な状態で維持されていました。これが、戦後の燃料転換(薪炭から石油へ)、肥料転換(堆肥から化学肥料へ)により、里山管理が衰退し、マツ林は荒廃。これがマツノザイセンチュウ被害の温床となりました。

これに対し、近年は里山再生の取り組みが各地で展開されています。(1)森林ボランティア活動、(2)里山保全条例、(3)森林経営計画への伝統知識の組み込み、(4)地域マツタケ生産との連動、により、伝統的な里山管理の現代的再生が進められています。

クロマツ防潮林の津波被害と再生

2011年東日本大震災では、東北沿岸のクロマツ防潮林が津波により壊滅的被害を受けました。代表的な被害として、(1)高田松原(岩手県陸前高田市):約7万本のクロマツが約1本(後の「奇跡の一本松」)を残し全壊。(2)仙台湾沿岸防潮林:宮城県沿岸約100km、津波で壊滅。(3)福島沿岸:原発事故による立入制限と防潮林被害の重複。被害総面積は約3,500haに及び、累積でクロマツ防潮林の30%が失われました。

これに対し、復興プロジェクトとして、(A)海岸林再生事業(2012年〜)、(B)盛土造成と新規植林、(C)抵抗性クロマツ品種の活用、(D)ボランティア参加型植林、(E)地域住民との協働、により、年間約500ha規模の海岸林再生が進められています。2024年時点で、約1.2万haが再生・成長期に入り、防潮機能の段階的回復が見られます。これは、自然災害と病害虫の複合被害から、地域の象徴的景観を取り戻す挑戦です。

まとめ:松枯れとの長期戦

マツ枯れは、(1)外来病害虫の侵入、(2)気候変動、(3)伝統的森林管理の衰退、(4)生態系劣化、(5)文化的景観の喪失、という多面的な課題を抱えた、日本最大の森林病害です。1980年ピーク240万m³から2023年34,000m³への縮小は、40年間の防除努力の成果ですが、(A)北海道への北上拡大、(B)抵抗性品種の普及、(C)海岸防潮林の再生、(D)伝統的里山管理の復活、(E)気候変動への適応、という新たな段階の課題が残されています。アカマツ・クロマツは日本の森林文化・景観の象徴であり、これらを次世代に継承するためには、科学的防除・伝統知識・地域協働・国際協力を組み合わせた、これからも長期的な総合戦略が、林業・地域・国家全体の重要な課題です。

関連記事

主要出典:林野庁「松くい虫被害量等」(各年度)、林野庁「森林・林業白書」、森林総合研究所「松くい虫被害対策技術」、各都道府県林業試験場、IPCC AR6 Working Group II(気候変動と森林病害虫)、日本林学会「マツ材線虫病」研究論文集。

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